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Specifically, species with larger, brighter, and more saturated color patches were found in habitats where the brightness and chroma of light were lower. Furthermore, environments with higher substrate temperature and higher relative humidity harbored species that produced secretions with a higher percentage of saturated and aromatic compounds. Thus, the elaboration of multimodal signals in this group of Cnemaspis geckos seems to increase the efficiency of communication in the signaling-environment, but the strength of signals in differe
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