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The PCA showed that ecological strategies of bryophytes in response to levels of irradiation were specialized in branch dwellers, although those of ground and trunk dwellers were less distinct. Conclusions Environmental filtering shaped the combination of functional traits and the spatial distribution of bryophytes along the vertical gradients. Bryophyte species from the upper canopy of cloud forests show narrow variation in functional traits in high-light intensity, whereas species in the lower vertical strata associated with low-light
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