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The genotype-phenotype relationship and the extensive knowledge on HPS3 function off their species strongly suggests that HPS3c.2420GA is the causative variation for the observed brown coat shade in French Bulldogs. To be able to obviously distinguish HPS3-related through the TYRP1-related brown coat color, and in range using the murine nomenclature, we suggest to designate this dog phenotype as "cocoa", additionally the mutant allele as HPS3co.the amount of Inter
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