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Harrison's rule, that body size is positively correlated between parasites and hosts, has been reported in a range of taxa, but whether the rule is applicable to cleptoparasitic insects is poorly understood. Subfamily Nomadinae, the largest group of cleptoparasitic bees, usurp the nests of a variety of host bees. Within the subfamily, Nomada exploits the most diverse hosts, using at least ten genera from five families. Here, we reassess the phylogeny of Nomadinae, including the expanded sampling of the genus Nomada, to explore host shif