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Environmental pH is a critical parameter for innumerable chemical reactions, myriad biological processes and all forms of life. The mechanisms that underlie the perception of external pH (pHe) have been elucidated in detail for bacteria, fungi and mammalian cells; however, little information is available on whether and, if so, how pHe is perceived by plants. This is particularly surprising since hydrogen ion activity of the substrate is of paramount significance for plants, governing the availability of mineral nutrients, the structure of
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