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Consequently, antioxidant enzymes such as catalase (CAT) superoxide dismutase (SOD) peroxidase (APX) and glutathione reductase (GR) activities were found to be significantly elevated in the desiccated state to avoid oxidative damage. Increased malondialdehyde (MDA) content and relative electrolyte leakage (REL) during desiccation provide evidence for membrane damage and loss of cell-wall integrity. During desiccation, the contents of osmolytes represented by sucrose and proline were found to increase to maintain cell structure integrity. After rehydration,