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The spiral ridge stores mechanical energy to decompress the pump when the muscle relaxes, and provides a sclerotized scaffold for the entire structure. The pumps do not occur in all Monotomidae, but seem to be restricted to the subfamily Monotominae. Two distinct forms were found, each in a different tribe (with one notable exception; Mimemodes having a sperm pump typical of Monotomini, not Europini). Moreover, the shapes of the pumps were found to be species-specific. The phylogenetic value of this structure is discussed, as sperm pump
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