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ciliata. Vapor pressure deficit (VPD) was linked with iWUE only in C. tabularis. Differences in shade tolerance, tree stature, and canopy position might have caused this species-specific variation in iWUE response to climate. Linear mixed effect modeling (LME) successfully simulated iWUE variability, explaining 41-51% of the total variance varying with species. Commonality analysis (CA) revealed that temperatures had a dominant influence on the inter-annual iWUE variability (64-77%) over precipitation (7-22%) and atmosphe