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We estimated male and female heritabilities for many characteristics known to be influenced by Y-chromosomes, including the amount of sternopleural bristles, abdominal bristles, intercourse brush teeth, and tibia length. A decrease in YN heritabilities weighed against Y1 would be in keeping with Y-chromosome variation being indication epistatic. A decrease in Y1 heritabilities is in keeping with Y-chromosome variation becoming additive, though additive-by-additive epistatic