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Fine root production accounts for a large proportion of net primary production (NPP) in forest ecosystems that is highly responsive to environmental and biotic changes. The underlying mechanisms of the relationship between tree species richness and fine root production have not been fully examined. Here we hypothesized that (i) the relationship between aboveground species richness and fine root production could be attributable to belowground spatial resource partitioning; (ii) either symmetrical or asymmetrical root proliferation to ob