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We find that risk factors associated with mortality differ between species, and that few generalizable patterns exist when bark beetle outbreaks occur in concert with a particularly long, hot drought. We see evidence that both long-term differences in physiology and shorter-term beetle-related selection and variability in defenses influence mortality susceptibility for ponderosa pine, whereas beetle dynamics may play a more prominent role in mortality patterns for white fir and pinyon pine. In contrast, incense cedar mortality appears
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