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The volatility of an environment significantly impacts cooperative behavior. In environments where viability-threatening events occur on a shorter timescale than reproduction, it is reasonable to measure the costs and benefits of cooperation in terms of their direct effect on survival probability. Then, the number of offspring increases with lifespan. With such a model, is it possible for cooperation to be evolutionarily stable, and how does cooperation depend on the benefit and cost of such interactions, and the volatility of the enviro
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