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We show that accounting for these two effects-variability in per-particle composition and deviations from the core-shell approximation-reconciles absorption enhancement predictions with laboratory and field observations and resolves the apparent discrepancy. Furthermore, our consistent model framework provides a path forward for improving predictions of BC's radiative effect on climate. Copyright © 2020 the Author(s). Published by PNAS.Developing lymphocytes diversify their antigen receptor (AgR) loci by variable (diversity) joining