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We also provide a more computationally efficient estimator of the asymptotic variance based on the non-parametric bootstrap. We evaluate the performance of our method using simulation and by application to an empirical dataset for gibbons. The method has been implemented in the PAUP* program, freely available at https//paup.phylosolutions.com for Macintosh, Windows and Linux operating systems. Supplementary data are available at Bioinformatics online. Supplementary data are available at Bioinformatics online.Uroplakin II (UPII) has be

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