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Understanding sexual segregation is crucial to comprehend sociality. A comparative analysis of long-term lion data from Serengeti and Ngorongoro in Tanzania, and Gir in India, reveals that male-female associations are contingent upon male and female group size, prey-size and availability, and the number of prides that each male coalition currently resides. Males maintain proximity with females, whereas females are responsible for segregation except at large kills. Lions feed on the largest prey in Ngorongoro and the smallest in Gir, and
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