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Pathogen dispersal characteristics, any fitness cost(s) of the resistance-breaking trait, the efficacy of host resistance, and the length of the timeframe of interest, all influence the strength of the spatial diversification effect. A key result is that spatial diversification has the strongest beneficial effect at intermediate fitness costs, an effect driven by a complex set of non-linear interactions. However, if the resistance-breaking strain cannot invade the landscape, then a partially effective resistance gene can result in spatial

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