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Moreover, the complex and context-specific regulation of Grem1, among a host of functionally convergent but structurally disparate BMP antagonists, warrants further research as we learn more about the pathogenic consequences of deranged expression of this small, but important, protein. © 2020 The Authors. The Journal of Pathology published by John Wiley Sons Ltd on behalf of Pathological Society of Great Britain and Ireland.1.An understudied aspect of vertebrate ecoimmunology has been the relative contributions of environmental fa