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This article is categorized under RNA Export and Localization RNA Localization RNA Evolution and Genomics Computational Analyses of RNA RNA Methods RNA Analyses in Cells. © 2020 The Authors. WIREs RNA published by Wiley Periodicals, Inc.The Waterloo Farm lagerstätte in South Africa provides a uniquely well-preserved record of a Latest Devonian estuarine ecosystem. Ecological evidence from it is reviewed, contextualised, and compared with that available from the analogous Swartvlei estuarine lake, with a particular emphasi