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Protected areas have been the cornerstone for conservation globally [1], but gaps still exist in preserving biodiversity [2]. Meanwhile, areas designated as protected have overlaps between designations and might vary in their management [3, 4]. All three phenomena-coverage gaps, overlapping designations, and disparities in management-are present in China [5, 6]. China plans to establish a national park system for the first time, aiming to reform the existing protected-area system [7-9]. However, there has been no quantitative spatial ana

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