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This study investigated the feasibility of combining natural ventilation and animal disturbance in constructed wetlands (CWs) and the joint effects on oxygen transfer, microbial activity, organics, and nitrogen removal. The results showed that natural ventilation extended the habitat depth of earthworms by approximately 10 cm by significantly improving oxygen transfer in CWs; in turn, the earthworms slightly promoted the addition of oxygen inside CWs through burrowing activity. Therefore, the interaction between natural ventilation and a

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