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A recent study resolved the recently sampled Lepisorus jakonensis as sister towards the staying genera in Lepisoreae excluding Paragramma, in addition to authors lumped all of the six well known genera into Lepisorus. In today's research, to infer a phylogeny we utilized DNA sequences of five plastid markers (matK, rbcL, rbcL-atpB, rps4 rps4-trnS, trnL trnL-F) of 172 accessions representing ca. 44 types of non-Lepisorus genera and 54 accessions representing ca. 50 types of Lepisorus as

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