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That arbuscular mycorrhizal (AM) fungi covary with plant communities is clear, and many papers report non-random associations between symbiotic partners. However, these studies do not test the causal relationship, or "codependency", whereby the composition of one guild affects the composition of the other. Here we outline underlying requirements for codependency, compare important drivers for both plant and AM fungal communities, and assess how host preference-a pre-requisite for codependency-changes across spatiotemporal scales and taxo

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