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On the other hand, our findings suggest that the use of insecticides with high toxicity is a necessary cause of the sharp population declines, as the declines did not occur when simulated toxicity levels were lower than those actually used. Overall, our numerical simulations suggest that the sharp population declines of S. frequens around the late 1990s were caused by the combined effects of insecticide use and farmland consolidation, in which rice paddies are converted to well-drained paddy fields. Conservation planning for dragonflies
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