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uction Regulation (CX2017-2), the Programme of Shanghai Academic/Technology Research Leader (17XD140090 and the Key Project of Shanghai Basic Research from Shanghai Municipal Science and Technology Commission (STCSM; 12JC140160. None of the authors has any conflict of interest to declare.Environmental variation experienced by a species across space and time can promote the maintenance of genetic diversity that may be adaptive in future global change conditions. Selection experiments have shown that purple sea urchin, Strongylocent