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nd evidence of an effect against more complex bacterial colonies.Biological forms occupy a constrained portion of theoretical morphospaces. Developmental models accounting for empirical morphospaces are necessary to achieve a better understanding of this phenomenon. We analyzed the phalangeal formulas (PFs) in lizards and relatives' hands by comparing them with a set of simulated PFs that compose a theoretical morphospace. We detected that (1) the empirical morphospace is severely limited in size, (2) the PFs comply with two properties