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Finally, we used the Mitani-Rodman defendability index to assess territoriality in P. nemaeus. We found that our population of P. nemaeus had a home range of 36.00 ha with a core area of 8.02 ha and an average daily path length of 509.11 ± 244.87 m. Daily path length was correlated to precipitation such that daily path lengths were shorter during increases in rainfall (r(17) = 0.52, p = .03). These measures resulted in a defendability index of 0.75, suggesting that the population of P. nemaus on Son Tra does not travel far enough each day