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Diversity in metacommunities is traditionally viewed to consist of the diversity within communities ([Formula see text]) that is complemented by the differences between communities ([Formula see text]) so as to result in the total diversity ([Formula see text]) of the metacommunity. This perception of the partitioning of diversity, where [Formula see text] is a function of [Formula see text] and [Formula see text] (usually [Formula see text] with all components specified as effective numbers), has several drawbacks, among which are (1) [F
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