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[This corrects the article DOI 10.1371/journal.pone.0230703.].Ecological theory implies that generalist species should have faculties with numerous adaptive peaks. Consequently, in heterogeneous environments such transformative landscapes can result in phenotypic divergence that becomes fixed in communities via reproductive isolation, thus driving speciation. Nevertheless, contrary to this expectation, the process of environmental diversification in wild communities is certainly not always connected w