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Trait association analysis of DNA markers independently developed from the Ma6 and Ma4 regions confirmed the mapping of Ma6 and Ma4. In the background of MaMa, 20.6% of acidity variation could be explained by Ma6, 28.5% by Ma4, and 50.7% by the combination of both. The effects of Ma6 and Ma4 in the background of Mama were also significant, but lower. These findings provide important genetic insight into high acidity in apple.Self-incompatibility (SI) to self-compatibility (SC) transition is one of the most frequent and prevalent evolutio
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