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4°C) lasted for 7 months. In winter, there were two melt periods. Vitality staining protocol showed a high number of viable (77.4% and 53.8%) and dormant cells (1.7% and 4.1%; capable of growth and reproduction once suitable conditions return) in the winter seepage and stream, respectively. NPQ and OJIP chlorophyll fluorescence parameters revealed several hours recovery of photosynthesis (both field and control samples). During recovery, only minor or mild stress on photosynthesis was detected. FV /FM values (the photosynthetic ef