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In turn, H. crenulatus had the highest contents of MUFAs and PUFAs in its gills and muscle tissues, including an important amount of eicosapentaenoic acid (EPA). The FA content of C. cinereus may indicate an adaptive physiological response aimed at maintaining its cellular fluid balance during periods of desiccation in the upper intertidal zone. In contrast, the FAs found in H. crenulatus may be linked to the high activity of the sodium‑potassium pump in its gills, in order to maintain osmoregulation in estuaries.In response to the conti